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NADPH oxidases generate ROS as O 2· − or H 2 O 2 depending on the isoform and context—NOX1/2/3 primarily release O 2· − that dismutates to H 2 O 2 , NOX4 predominantly yields H 2 O 2 and Ca²⁺-activated NOX5 together with epithelial DUOX1/2 contribute directly to H 2 O 2 production at barrier surfaces (Kim and Moon 2024)